Planthoppers: FLOW Website

Guianaphrynini Gnezdilov, 2018

Cordelini Gnezdilov, 2019

Issinae Spinola, 1839

Hemisphaeriinae Melichar, 1906

Inoderbidae† Shcherbakov & Emeljanov, 2021

Jubisentidae† Zhang, Ren & Yao, 2019

Issidae Spinola, 1839

Thioniinae Melichar, 1906

Thioniini Melichar, 1906

Tribe

Thioniini Melichar, 1906

Basionym original publication

Melichar L. 1906 - Monographie der Issiden. (Homoptera). Abhandlungen der K. K. Zoologisch-botanischen Gesellschaft in Wien. Wien 3: 1-327 [3].

Chronological account

Thioniini Melichar, 1906 synonym of Issini Spinola, 1839 according to Gnezdilov (2013): 725

Thioniina Melichar, 1906 [Fulgoromorpha, Fulgoroidea, Issidae, Issinae, Issini] status revivisco previous rank of Thioniini Melichar, 1906 [Fulgoromorpha, Fulgoroidea, Issidae, Thioniinae] according to Wang et al. (2016): 232

Thioniini Melichar, 1906 [Fulgoromorpha, Fulgoroidea, Issidae, Issinae] status revivisco previous rank of Thioniina Melichar, 1906 [Fulgoromorpha, Fulgoroidea, Issidae, Issinae, Issini] according to Gnezdilov (2016): 362

Thioniini Melichar, 1906 - Graphical schematical display

Caption


Valid name	Historical link	Change to lower rank

Basionym	Synonymy	Change to upper rank

	Homonymys

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116 taxa (97 species, 14 genera)

> Amnisa Stål, 1862

Amnisa lata Schmidt, 1910

Amnisa singularis Stål, 1862 type species

Amnisa verticalis Melichar, 1906

> Diceroptera Gnezdilov, 2011

Diceroptera humboldti Gnezdilov, 2011 type species

> Dracela Signoret, 1861

Dracela acuta (Metcalf, 1938)

Dracela annulipes Signoret, 1861 type species

Dracela pehlkei Schmidt, 1923

> Heremon Kirkaldy, 1903

Heremon cribatum (Melichar, 1906)

Heremon gibbifrons Melichar, 1906

Heremon infixum (Walker, 1858)

Heremon instabile (Stål, 1862)

Heremon notatum (Melichar, 1906)

> Oronoquina Gnezdilov, 2018

> Oronoqua Fennah, 1947 type genus

Oronoqua deina Fennah, 1947 type species

Oronoqua feria Gnezdilov & Bartlett, 2020

Oronoqua ibisca Gnezdilov, Bonfils, Aberlenc & Basset, 2010

Oronoqua orellana Gnezdilov & Bartlett, 2020

> Paranipeus Melichar, 1906

Paranipeus latipes Melichar, 1906 type species

> Proteinissus Fowler, 1904

Proteinissus bilimeki Fowler, 1904 type species

Proteinissus isolatus (Caldwell, 1945)

Proteinissus pentagonatus (Caldwell, 1945)

Proteinissus punctatus (Caldwell, 1945)

Proteinissus reticulatus (Caldwell, 1945)

Proteinissus viridus (Caldwell, 1945)

> Thioniamorpha Metcalf, 1938

Thioniamorpha marmorata Metcalf, 1938

> Thioniella Metcalf, 1938

Thioniella rugosa Metcalf, 1938

> Thioniina Melichar, 1906

> Aplos Gnezdilov, 2018

Aplos simplex (Germar, 1830) type species

> Cheiloceps Uhler, 1895

Cheiloceps anguillana (Fennah, 1965)

Cheiloceps argo (Fennah, 1949)

Cheiloceps borinqueta (Caldwell & Martorell, 1951)

> Cheiloceps musca Uhler, 1895 type species

Thionia (Cheiloceps) musca gredanada Fennah, 1955

Cheiloceps puertoricensis (Caldwell & Martorell, 1951)

Thionia (Cheiloceps) clusiae Fennah, 1955

Thionia (Cheiloceps) laodice Fennah, 1955

Thionia (Cheiloceps) medusa Fennah, 1955

> Fowlerium Gnezdilov, 2018

Fowlerium acutum (Doering, 1941)

Fowlerium naso (Fowler, 1904) type species

Fowlerium productum (Van Duzee, 1908)

> Thionia Stål, 1859 type genus

Thionia biforis (Germar, 1830)

Thionia boliviensis Schmidt, 1910

Thionia borinquensis Dozier, 1931

Thionia brasiliensis Schmidt, 1910

Thionia brevifrons Schmidt, 1910

Thionia brevior Fowler, 1904

Thionia bufo Fennah, 1945

> Thionia bullata (Say, 1830)

Thionia bullata bullata (Metcalf, 1938)

Thionia carinata Melichar, 1906

Thionia caviceps Fowler, 1905

Thionia coriacea (Fabricius, 1803)

Thionia crucifera Metcalf, 1938

Thionia dissimilis Schmidt, 1910

Thionia douglundbergi† Stroinski & Szwedo, 2008

Thionia dryas Fennah, 1945

Thionia dubiosa Melichar, 1906

Thionia ecuadoriensis Schmidt, 1910

Thionia elliptica (Germar, 1830)

Thionia fowleri Metcalf, 1938

Thionia gibba Melichar, 1906

Thionia gibbicollis Dozier, 1931

Thionia herbecea (Spinola, 1839)

Thionia humilis Fowler, 1904

Thionia impressa Melichar, 1906

Thionia latifrons Melichar, 1906

Thionia longipennis (Spinola, 1839) type species

Thionia maculata Melichar, 1906

Thionia maculipes Stål, 1864

Thionia mammifera Fennah, 1945

Thionia mexicana Melichar, 1906

Thionia minor Schmidt, 1910

Thionia obrienae Wilson, 1987

Thionia obsoleta Melichar, 1906

Thionia ocellata Melichar, 1906

Thionia ohausi Schmidt, 1910

Thionia omani Doering, 1939

Thionia onerata Melichar, 1906

Thionia ovata Melichar, 1906

Thionia parana Bergroth, 1910

Thionia pehlkei Schmidt, 1910

Thionia pictifrons Fowler, 1905

Thionia prasina (Spinola, 1839)

Thionia quadratifrons Schmidt, 1910

Thionia quinquata Metcalf, 1923

Thionia ramosi Caldwell & Martorell, 1951

Thionia rubrocostata (Spinola, 1839)

Thionia schmidti Schmidt, 1910

Thionia scutellata Fowler, 1904

Thionia similis Schmidt, 1910

Thionia soluta Fowler, 1905

Thionia sordida Fowler, 1904

Thionia stipes Fowler, 1905

Thionia tigrata Melichar, 1906

Thionia transversalis Melichar, 1906

Thionia ustulipunctata (Uhler, 1876)

Thionia variata Melichar, 1906

Thionia variegata Stål, 1864

> Waoraniina Gnezdilov & Bartlett, 2018

> Waorania Gnezdilov & Bartlett, 2018 type genus

Waorania jaguarina Gnezdilov & Bartlett, 2018 type species

Waorania pantherina Gnezdilov & Bartlett, 2018 type species

Distribution map: extant taxa

Opacity 30%

Data accuracy

Level 4

Levels 1, 2 and 3

(TDWG standard)

GBIF occurrences

Geographical distribution

Austria according to Holzinger et al. (2020): 11

Bolivia according to Melichar (1906): 275, Melichar (1906): 272, Schmidt (1910): 196, Schmidt (1910): 193, Schmidt (1910): 205

Brazil according to Germar (1830): 51, Spinola (1839): 357, Spinola (1839): 349, Stål (1854): 247, Melichar (1906): 275, Melichar (1906): 273, Melichar (1906): 271, Schmidt (1910): 202, Schmidt (1910): 201, Metcalf (1938): 410, Metcalf (1945): 136

Brazil North (Amazonas) according to Melichar (1906): 274, Melichar (1906): 280, Melichar (1906): 278

Brazil Northeast (Bahia) according to Melichar (1906): 275, Schmidt (1910): 201

Brazil North (Pará) according to Melichar (1906): 274, Melichar (1906): 277, Metcalf (1938): 410

Brazil North (Rondônia) according to Gnezdilov & O'Brien (2008): 19

Brazil Southeast (Espirito Santo) according to Melichar (1906): 278

Brazil Southeast (Rio de Janeiro) according to Walker (1858): 88, Stål (1862): 10, Stål (1862): 9, Melichar (1906): 275, Melichar (1906): 273, Melichar (1906): 258, Melichar (1906): 271, Melichar (1906): 261, Schmidt (1910): 202

Brazil Southeast (São Paulo) according to Melichar (1906): 258, Melichar (1906): 271

Brazil South (Rio Grande do Sul) according to Melichar (1906): 258

Brazil South (Santa Catarina) according to Schmidt (1910): 183, Schmidt (1910): 190

Canada (Ontario) according to Bartlett et al. (2014): 172

Colombia according to Melichar (1906): 274, Melichar (1906): 273, Melichar (1906): 226, Melichar (1906): 271, Schmidt (1910): 206, Schmidt (1910): 198, Schmidt (1923): 18

, Gnezdilov & O'Brien (2008): 21

Costa Rica according to Schmidt (1910): 207, Metcalf (1938), Gnezdilov & O'Brien (2008): 19

Cuba according to Uhler (1876): 355

Democratic Republic of the Congo according to Synave (1973)

Dominica according to Gibson (1919): 277, Fennah (1955): 36, Gnezdilov (2018): 162

Dominican Republic according to Stroinski & Szwedo (2008)

Ecuador according to Schmidt (1910): 203, Schmidt (1910): 199, Gnezdilov & O'Brien (2008): 19, Gnezdilov & Bartlett (2018): 69

, Gnezdilov & Bartlett (2018): 68

, Gnezdilov & Bartlett (2020): 69

Florida according to Bartlett et al. (2014): 172

France (French Guiana) according to Signoret (1861): 501, Melichar (1906): 260, Melichar (1906): 226, Melichar (1906): 280, Gnezdilov et al. (2010)

Great Britain (Anguilla) according to Fennah (1965): 105

Great Britain (British Virgin Islands) according to Fennah (1949): 60

, Gnezdilov (2018): 163

Great Britain (Montserrat) according to Fennah (1955): 37

Grenada according to Fennah (1955): 34

Guatemala according to Fowler (1905): 125, Melichar (1906): 272

Guyana according to Metcalf (1945): 136, Fennah (1947)

Haiti according to Dozier (1931): 17

Honduras according to Schmidt (1910): 191

Italy according to Gnezdilov & Poggi (2014): 240

, Gnezdilov (2018): 166

Jamaica according to Melichar (1906): 284

Mexico according to Stål (1864): 51, Uhler (1876): 355, Melichar (1906): 285

Mexico Central (Mexico Distrito Federal) according to Caldwell (1945): 111, Caldwell (1945): 109, Gnezdilov & O'Brien (2008): 19

Mexico Central (México State) according to Caldwell (1945): 111, Gnezdilov & O'Brien (2008): 19

Mexico Central (Morelos) according to Fowler (1904): 121, Caldwell (1945): 109, Gnezdilov & O'Brien (2008): 19

Mexico Central (Puebla) according to Caldwell (1945): 112, Caldwell (1945): 111, Gnezdilov & O'Brien (2008): 19, Gnezdilov (2018): 164

Mexico Gulf (Veracruz) according to Fowler (1904): 123, Fowler (1904): 124, Melichar (1906): 272, Herrera (1923): 116

Mexico Northeast (Hidalgo) according to Caldwell (1945): 118, Bartlett et al. (2014): 172, Gnezdilov (2018): 164

Mexico Southeast (Chiapas) according to Bourgoin et al. (2020): 2

Mexico Southwest (Guerrero) according to Fowler (1904): 124, Fowler (1905): 125

Mexico Southwest (Jalisco) according to Fowler (1904): 121, Caldwell (1945): 109

Nicaragua according to Melichar (1906): 281, Metcalf (1938): 409

Panama according to Fowler (1904): 123, Fowler (1905): 126, Fowler (1905): 125, Fowler (1905): 127, Melichar (1906): 272, Metcalf (1938): 413, Metcalf (1938): 410, Metcalf (1938): 414, Metcalf (1938): 411, Metcalf (1938): 412, Metcalf (1945): 136, Gnezdilov & O'Brien (2008): 19, Gnezdilov et al. (2010)

Peru according to Melichar (1906): 276, Melichar (1906): 272, Gnezdilov & O'Brien (2008): 19

Saint Lucia according to Fennah (1955): 35, Gnezdilov (2018): 162

Saint Vincent and the Grenadines according to Uhler (1895): 68

Trinidad-Tobago according to Fennah (1945): 508, Fennah (1945): 509

United States of America (Alabama) according to Benton & McCreadie (2009), Bartlett et al. (2014): 172, Bartlett et al. (2014): 173

United States of America (Arizona) according to Bartlett et al. (2014): 172

United States of America (Arkansas) according to O'Brien (1988): 868, Bartlett et al. (2014): 173, Bartlett et al. (2014): 172

United States of America (Colorado) according to Van Duzee (1908): 494, Bartlett et al. (2014): 172

United States of America (Delaware) according to Bartlett et al. (2014): 172, Bartlett et al. (2014): 173

United States of America (District of Columbia) according to Bartlett et al. (2014): 173, Bartlett et al. (2014): 172

United States of America (Florida, nearctic) according to Bartlett et al. (2014): 173

United States of America (Georgia) according to Bartlett et al. (2014): 173, Bartlett et al. (2014): 172

United States of America (Illinois) according to Bartlett et al. (2014): 173, Bartlett et al. (2014): 172

United States of America (Indiana)

United States of America (Kentucky) according to Germar (1830): 51, Germar (1830), O'Brien (1988): 868, Bartlett et al. (2014): 172, Bartlett et al. (2014): 173

United States of America (Louisiana) according to Bartlett et al. (2014): 172

United States of America (Maryland) according to Bartlett et al. (2014): 172, Bartlett et al. (2014): 173

United States of America (Mississippi) according to Bartlett et al. (2014): 172, Bartlett et al. (2014): 173

United States of America (Missouri) according to O'Brien (1988): 868, Bartlett et al. (2014): 173, Bartlett et al. (2014): 172

United States of America (New Jersey) according to Bartlett et al. (2014): 172, Bartlett et al. (2014): 173

United States of America (New Mexico) according to Doering (1939): 461, Bartlett et al. (2014): 172

United States of America (New York) according to Bartlett et al. (2014): 172

United States of America (North Carolina) according to O'Brien (1988): 868, Bartlett et al. (2014): 172, Bartlett et al. (2014): 173

United States of America (Ohio) according to Bartlett et al. (2014): 172, Bartlett et al. (2014): 173

United States of America (Oklahoma) according to Bartlett et al. (2014): 172

United States of America (Pennsylvania) according to O'Brien (1988): 868, Bartlett et al. (2014): 173, Bartlett et al. (2014): 172

United States of America (Puerto Rico) according to Dozier (1931): 18, Caldwell & Martorell (1951): 252, Caldwell & Martorell (1951): 249, Caldwell & Martorell (1951): 249, Caldwell & Martorell (1951): 251

United States of America (South Carolina) according to Bartlett et al. (2014): 173, Bartlett et al. (2014): 172

United States of America (Tennessee) according to Bartlett et al. (2014): 173, Bartlett et al. (2014): 172

United States of America (Texas) according to Doering (1941): 214, O'Brien (1988): 868, Bartlett et al. (2014): 172, Bartlett et al. (2014): 173, Gnezdilov (2018): 165

United States of America (Utah) according to Bartlett et al. (2014): 172

United States of America (Virginia) according to Bartlett et al. (2014): 172, Bartlett et al. (2014): 173

United States of America (Virgin Islands) according to Caldwell & Martorell (1951): 249

Venezuela according to Lethierry (1890): 149, Melichar (1906), Gnezdilov (2011): 120

Biological interactions

Host-plant

Alnus jorullensis Kunth (Fagales, Betulaceae) according to Gnezdilov (2018): 164

Clusia sp. (Theales, Clusiaceae) according to Fennah (1955): 35, Gnezdilov (2018): 162

Coccoloba uvifera (L.) L. (Caryophyllales, Polygonaceae) according to Llano-Arias et al. (2022): 7

Hedera helix L. (Apiales, Araliaceae) according to Holzinger et al. (2020): 11

Juniperus monosperma (Engelm.) Sarg. (Cupressales, Cupressaceae) according to Gnezdilov (2018): 165

Pinus taeda L. (Pinales, Pinaceae) according to Llano-Arias et al. (2022): 7

Pinus virginiana Mill. (Pinales, Pinaceae) according to Llano-Arias et al. (2022): 7

Quercus ilicifolia Wangenh. (Fagales, Fagaceae) according to Llano-Arias et al. (2022): 7

Quercus marilandica Münchh. (Fagales, Fagaceae) according to Llano-Arias et al. (2022): 7

Ulmus alata Michx (Rosales, Ulmaceae) according to Wheeler & Wilson (1988)

Ulmus laevis Pall. (Rosales, Ulmaceae) according to Gnezdilov & Poggi (2014): 240

Pinus echinata Mill. (Pinales, Pinaceae) according to Llano-Arias et al. (2022): 7


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