Planthoppers: FLOW Website

. Lophopidae incertae sedis

Menoscinae Melichar, 1915

Lalacidae† Hamilton, 1990

Meenoplidae Fieber, 1872

Delphacoidea Leach, 1815

Surijokocixioidea† Shcherbakov, 2000

Fulgoroidea Latreille, 1807

Lophopidae Stål, 1866

Lophopinae Stål, 1866

Lophopini Stål, 1866

Tribe

Lophopini Stål, 1866

Basionym original publication

Stål C. 1866 - Flatidae. Hemiptera Africana 4: 235-251 [130].

104 taxa (73 species, 16 genera)

> Acothrura Melichar, 1915

Acothrura fucizona Wang, Chou & Yuan, 2000

Acothrura impunctata (Jacobi, 1905) type species

Acothrura orientalis (Liang, 2000)

> Asantorga Melichar, 1915

Asantorga tuberculata Melichar, 1915 type species

> Corethrura Hope, 1843

Corethrura fuscovaria Hope, 1843 type species

> Jivatma Distant, 1906

Jivatma metallica Distant, 1906 type species

> Katoma Baker, 1925

Katoma trichurensis Baker, 1925

> Lacusa Stål, 1862

Lacusa digitata Xing & Chen, 2014

Lacusa fuscofasciata (Stål, 1854) type species

Lacusa producta Xing & Chen, 2014

Lacusa yunnanensis Chou & Huang, 1985

> Lophops Spinola, 1839 type genus

Lophops africana (Schmidt, 1912)

Lophops angustipennis Stål, 1866

Lophops bwamba Soulier-Perkins, 2017

Lophops carinata (Kirby, 1891)

Lophops fusca Melichar, 1915

Lophops incognita Soulier-Perkins, 2017

Lophops intermedia Schmidt, 1926

Lophops kwazulu Soulier-Perkins, 2017

Lophops malagarassi Soulier-Perkins, 2017

Lophops mentawiensis Baker, 1927

Lophops pallida Melichar, 1902

Lophops saccharicida (Kirkaldy, 1906)

Lophops servillei Spinola, 1839 type species

Lophops sigwalti Soulier-Perkins, 2017

Lophops stilleri Soulier-Perkins, 2017

Lophops tigris Fennah, 1955

Lophops verschureni Synave, 1962

Lophops watshami Soulier-Perkins, 2017

Lophops zebra (Walker, 1870)

> Maracota Emeljanov, 2018

Maracota soulierae Emeljanov, 2018 type species

> Paracorethrura Melichar, 1915

Paracorethrura arrhen Soulier-Perkins & Stroinski, 2017

Paracorethrura iocnemis (Jacobi, 1905) type species

> Pitambara Distant, 1906

Pitambara acuminata Lv, Yang, Zhang & Chen, 2025

Pitambara assamensis Liang, 2000

Pitambara borneensis Baker, 1925

Pitambara dawnana Distant, 1912

Pitambara impudica Wang & Bourgoin, 2016

Pitambara interrupta Distant, 1906

Pitambara montana Distant, 1912

Pitambara nigrofuscata Liang, 2000

Pitambara radians (Kirby, 1891) type species

Pitambara rubrofasciata Liang, 2000

Pitambara ruiliensis Lv, Yang, Zhang & Chen, 2025

Pitambara sinuata Distant, 1906

Pitambara tagalica (Stål, 1854)

Pitambara tonkinensis Lallemand, 1942

Pitambara tricorne Wang & Wang, 2016

Pitambara triremiprocta Wang & Soulier-Perkins, 2016

Pitambara trypetoides Emeljanov, 2018

Pitambara undulata Distant, 1906

> Podoschtroumpfa Soulier-Perkins, 1998

Podoschtroumpfa magna Soulier-Perkins, 1998 type species

Podoschtroumpfa rubrolineata Liang, 2000

> Pyrilla Stål, 1859

> Pyrilla aberrans (Kirby, 1891)

Pyrilla aberrans aberrans Kirby, 1891

Pyrilla aberrans achates Fennah, 1963

Pyrilla aberrans comes Fennah, 1963

Pyrilla aberrans consors Fennah, 1963

Pyrilla aberrans palghati Fennah, 1963

Pyrilla lycoides (Walker, 1862)

> Pyrilla perpusilla (Walker, 1851)

Pyrilla perpusilla chikkaballapurana Fennah, 1963

Pyrilla perpusilla coimbatorensis Fennah, 1963

Pyrilla perpusilla dhimbami Fennah, 1963

Pyrilla perpusilla lycoides Walker, 1862

Pyrilla perpusilla naraikkaduana Fennah, 1963

Pyrilla perpusilla perpusilla (Walker, 1851)

Pyrilla perpusilla pirmedana Fennah, 1963

Pyrilla perpusilla pusana Distant, 1914

Pyrilla perpusilla singhalensis Fennah, 1963

Pyrilla perpusilla thekkadiana Fennah, 1963

Pyrilla protuberans Stål, 1859 type species

Pyrilla punjabensis Shakila, 1984

Pyrilla rahimyarensis Shakila, 1984

Pyrilla sinica Liang, 1997

Pyrilla sumatrensis Baker, 1925

> Sarebasa Distant, 1909

Sarebasa celebris Distant, 1909 type species

> Sarmatoca Emeljanov, 2018

Sarmatoca cathemerina Emeljanov, 2018

> Serida Walker, 1857

Serida balteata Distant, 1909

Serida castanea Emeljanov, 2018

Serida elasmoscelis (Jacobi, 1944)

Serida excisa Melichar, 1915

Serida gorgopis Fennah, 1978

Serida latens Walker, 1857 type species

Serida parenthesisflexuosa Wang & Soulier-Perkins, 2016

Serida shcherbakovi Emeljanov, 2018

> Silvispina Wang & Soulier-Perkins, 2016

Silvispina changpotou Wang & Soulier-Perkins, 2016 type species

Distribution map: extant taxa

Opacity 30%

Data accuracy

Level 4

Levels 1, 2 and 3

(TDWG standard)

GBIF occurrences

Geographical distribution

Afghanistan according to Dlabola (1957): 274

Africa according to Soulier-Perkins & Wilson (2017): 483

Algeria according to Soulier-Perkins & Wilson (2017): 482

Bangladesh according to Hope (1843): 135

, Distant (1906): 243

Benin according to Soulier-Perkins & Wilson (2017): 487

Cambodia according to Liang (1997), Soulier-Perkins (1998): 613, Liang (2000): 301

Cameroon according to Gerstaecker (1895): 17, Melichar (1915), Van Stalle (1982): 17

, Soulier-Perkins & Wilson (2017): 481

Cape Provinces (Eastern Cape) according to Soulier-Perkins & Wilson (2017): 484

Cape Provinces (Western Cape Province) according to Soulier-Perkins & Wilson (2017): 486

Central African Republic according to Soulier-Perkins & Wilson (2017): 487

Chad according to Soulier-Perkins & Wilson (2017): 487

China (Fujian) according to Liang (1996): 148 refuted by Liang (1997): 36, Liang (1997): 36, Wang et al. (2016): 2

China (Guangdong) according to Fennah (1956): 521, Fennah (1956): 520, Liang (1996): 147 refuted by Wang et al. (2016): 16

, Liang (2000): 288, Xing & Chen (2014): 142

, Wang et al. (2016): 2

China (Guangxi) according to Jacobi (1905), Liang (1996): 148 refuted by Liang (1997): 38, Liang (1997): 38, Wang et al. (2016): 2

China (Hainan) according to Fennah (1956): 521, Fennah (1956): 520, Liang (1996): 148, Liang (1997): 38, Liang (2000): 288, Xing & Chen (2014): 142

, Wang et al. (2016): 2

China (Jiangxi) according to Liang (1996): 148 refuted by Liang (1997): 37, Liang (1997): 37, Wang et al. (2016): 2

China South-Central (Guizhou) according to Liang (1996): 147 refuted by Wang et al. (2016): 15

, Liang (2000): 288, Xing & Chen (2014): 142

, Wang et al. (2016): 2

China (Yunnan) according to Jacobi (1905), Chou et al. (1985): 128, Liang (1996): 147 refuted by Wang et al. (2016): 16

, Liang (1996): 149, Liang (1997): 38, Liang (1997), Wang et al. (2000): 246, Liang (2000): 291, Liang (2000): 288, Wang et al. (2000), Liang (2000): 297, Liang (2005): 430, Wang & Wang (2012): 632, Xing & Chen (2014): 145

, Xing & Chen (2014): 142

, Xing & Chen (2014): 147

, Wang et al. (2016): 14

, Wang et al. (2016): 2

, Wang et al. (2016): 8

, Wang et al. (2016): 10

, Wang & Soulier-Perkins (2016): 238

, Wang et al. (2016): 6

, Wang et al. (2016): 21

, Wang et al. (2016): 23

, Lv et al. (2025): -, Lv et al. (2025):

Democratic Republic of the Congo according to Soulier-Perkins & Wilson (2017): 481

, Soulier-Perkins & Wilson (2017): 491

Egypt according to Soulier-Perkins & Wilson (2017): 235

Equatorial Guinea according to Melichar (1915), Soulier-Perkins & Wilson (2017): 481

Ghana according to Soulier-Perkins & Wilson (2017): 487

, Soulier-Perkins & Wilson (2017): 482

Guinea according to Lallemand (1958): 225

, Soulier-Perkins & Wilson (2017): 481

, Soulier-Perkins & Wilson (2017): 487

India according to Xing & Chen (2014): 142

India (Assam) according to Liang (2000): 288, Liang (2000): 293, Liang (2000): 295

India (Bihar) according to Fennah (1963): 721

India (Chattisgarh) according to Sankararaman et al. (2020): 15927

India (Delhi) according to Kumar et al. (2015): 433

India (Gujarat) according to Sankararaman et al. (2020): 15927

India (Haryana) according to Sankararaman et al. (2020): 15927

India (Karnataka) according to Fennah (1963): 731, Fennah (1963): 723, Fennah (1963): 725

India (Kerala) according to Fennah (1963): 730, Fennah (1963): 726, Fennah (1963): 723

India (Maharashtra) according to Distant (1906): 243 , Sankararaman et al. (2020): 15927

India (Odisha, Orissa) according to Sankararaman et al. (2020): 15927

India (Punjab) according to Sankararaman et al. (2020): 15927

India (Rajasthan) according to Sankararaman et al. (2020): 15927

India (Tamil Nadu) according to Distant (1906): 243 , Fennah (1963): 722, Fennah (1963): 725, Fennah (1963): 730, Fennah (1963): 731

India (Uttar Pradesh) according to Fennah (1963): 721, Gangwar et al. (2008): 162, Sankararaman et al. (2020): 15927

India (West Bengal) according to Distant (1906): 243 , Fennah (1963): 720

Indonesia (Borneo) according to Distant (1906): 243

Indonesia (Jawa) according to Melichar (1914): 104, Fennah (1963): 732

Iran according to Mozaffarian & Wilson (2011): 35, Soulier-Perkins & Wilson (2017): 486

Ivory Coast according to Lallemand (1942): 68

, Soulier-Perkins & Wilson (2017): 487

Japan (Honshu) according to Hayashi & Fujinuma (2016): 355

Japan (Kyushu) according to Hayashi & Fujinuma (2016): 355

Japan (Shikoku) according to Hayashi & Fujinuma (2016): 355

Kenya according to Soulier-Perkins & Wilson (2017): 484

KwaZulu-Natal according to Soulier-Perkins & Wilson (2017): 484

Laos according to Liang (2000): 291, Wang et al. (2016): 23

Malaysia (peninsula) according to Liang (2000): 288

Malaysia (Sabah) according to Liang (2000): 288

Malaysia (Sarawak) according to Lallemand (1939): 71

Myanmar (ex Burma) according to Distant (1906): 243

, Distant (1912): 189, Xing & Chen (2014): 142

Nansei-shoto (Ryukyu Islands) according to Hayashi & Fujinuma (2016): 355

Nepal according to Neupane (1976)

, Liang (2000): 288, Xing & Chen (2014): 142

Nigeria according to Soulier-Perkins & Wilson (2017): 482

, Soulier-Perkins & Wilson (2017): 481

Oman according to Soulier-Perkins & Wilson (2017): 486

Pakistan (palearctic) according to Shakila (1984)

Saudi Arabia according to Soulier-Perkins & Wilson (2017): 486

Senegal according to Stål (1866): 202, Lallemand (1958): 225

, Soulier-Perkins & Wilson (2017): 487

, Soulier-Perkins & Wilson (2017): 489

, Soulier-Perkins & Wilson (2017): 482

Sierra Leone according to Stål (1855): 95

, Stål (1866): 202, Soulier-Perkins & Wilson (2017): 482

, Soulier-Perkins & Wilson (2017): 481

, Soulier-Perkins & Wilson (2017): 487

Solomon Islands (North Solomons) according to Fennah (1955): 170

Solomon Islands (South Solomons) according to Fennah (1955): 170

South Africa according to Stål (1866): 202

South Africa (Northern provinces, ex Transvaal) according to Soulier-Perkins & Wilson (2017): 491

South Africa-Northern Provinces (Gauteng) according to Soulier-Perkins & Wilson (2017): 491

Sri Lanka according to Kirby (1891): 140

, Fennah (1963): 727, Fennah (1963): 729

Sudan according to Soulier-Perkins & Wilson (2017): 482

Taiwan according to Wang et al. (2016): 2

, Hayashi & Fujinuma (2016): 355

Tanzania according to Soulier-Perkins & Wilson (2017): 485

Thailand according to Fennah (1963): 727, Liang (2000): 299, Liang (2000): 301, Liang (2000): 288, Xing & Chen (2014): 142

, Wang et al. (2016): 23

, Wang et al. (2016): 21

Togo according to Soulier-Perkins & Wilson (2017): 481

Tunisia according to Linnavuori (1973)

Uganda according to Soulier-Perkins & Wilson (2017): 482

United Arab Emirates according to Soulier-Perkins & Wilson (2017): 486

Vietnam according to Fennah (1978): 274, Fennah (1978): 275, Liang (2000): 291, Liang (2000): 301, Pham & Ta (2006): 521, Pham & Ta (2006): 522, Pham & Ta (2006): 520, Soulier-Perkins & Stroinski (2017)

, Emeljanov (2018): 461

, Emeljanov (2018): 460

, Emeljanov (2018): 454

, Emeljanov (2018): 459

, Emeljanov (2018): 456

, Emeljanov (2018): 457

Vietnam (Tonkin) according to Lallemand (1942): 74, Wang et al. (2016): 23 , Wang et al. (2016): 21 , Soulier-Perkins & Stroinski (2017)

West Himalaya (Uttaranchal = Uttarakhand) according to Sankararaman et al. (2020): 15927

Zimbabwe (ex Rhodesia) according to Soulier-Perkins & Wilson (2017): 492

Biological interactions

Host-plant

Bambusa sp. (Poales, Poaceae) according to Emeljanov (2018): 461

, Emeljanov (2018): 454

Gossypium sp. (Malvales, Malvaceae) according to Soulier-Perkins & Wilson (2017): 486

Oryza sativa L. (Poales, Poaceae) according to Shakila (1984)

Saccharum officinarum L. (Poales, Poaceae) according to Kirkaldy (1907): 123, Fennah (1963): 722, Fennah (1963): 721, Soulier-Perkins & Wilson (2017): 484

Saccharum ravennae (L.) L. (Poales, Poaceae) according to Soulier-Perkins & Wilson (2017): 486

Saccharum sp. (Poales, Poaceae) according to Shakila (1984), Shakila (1984), Shakila (1984), Shakila (1984), Shakila (1984), Shakila (1984)

Sorghum halepense (L.) Pers. (Poales, Poaceae) according to Gangwar et al. (2008): 163

Zea mays L. (Poales, Poaceae) according to Shakila (1984), Shakila (1984), Shakila (1984)

Parasitism: parasitoid

Aprostocetus gala (Walker, 1847) (Insecta, Hymenoptera, Eulophidae) according to Kumarasinghe & Wratten (1996): 489

Cheiloneurus pyrillae Mani, 1939 (Insecta, Hymenoptera, Encyrtidae) according to Kumarasinghe & Wratten (1996): 489

Dryinus indicus (Kieffer, 1914) (Insecta, Hymenoptera, Dryinidae) according to Mita (2009): 50

Dryinus pyrillae (Kieffer, 1911) (Insecta, Hymenoptera, Dryinidae) according to Kumarasinghe & Wratten (1996): 490

, Mita (2009): 51

Dryinus pyrillivorus Olmi, 1986 (Insecta, Hymenoptera, Dryinidae) according to Mita (2009): 51

Erythraeus (Zaracarus) perpusillus Kamran et al. 2009 (Acari, Trombidiformes, Erythraeidae) according to Kamran et al. (2009): 199, Stroinski et al. (2013): 199

Fulgoraecia melanoleuca (Fletcher, 1939) (Insecta, Lepidoptera, Epipyropidae) according to Kumarasinghe & Wratten (1996): 490

, Gangwar et al. (2008): 162, Kumar et al. (2015): 433

, Sankararaman et al. (2020): 15927

Halictophagus compactus (Pierce, 1914) (Insecta, Strepsiptera, Halictophagidae) according to Kumarasinghe & Wratten (1996): 489

Ooencyrtus pyrillae (Mani, 1939) (Insecta, Hymenoptera, Encyrtidae) according to Kumarasinghe & Wratten (1996): 489

Parachrysocharis javensis Girault, 1913 (Insecta, Hymenoptera, Eulophidae) according to Kumarasinghe & Wratten (1996): 489

Proleuroceroides pyrillae (Crawford, 1916) (Insecta, Hymenoptera, Encyrtidae) according to Kumarasinghe & Wratten (1996): 489

Tetrastichus pyrillae Crawford, 1916 = Parachrysocharis javensis Girault, 1913 (Insecta, Hymenoptera, Eulophidae) according to Swaminathan & Ananthakrishnan (1984): 8


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