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地质计年
Bartonian [41.2-37.8 Ma] 根 据 Brysz & Szwedo (2018): 4
Lutetian [47.8-41.2 Ma] 根 据 Lefebvre et al. (2007): 42
地理分布图: 現存類群
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地理分布
Poland  根 据  Lefebvre et al. (2007): 42